There are actually two stimuli present (you can add more) and each of them move toward a single direction. However, they alternate, so it looks like they're transparent. This alternating pattern is technically known as PWM. If you increase the PWM period (the length of each alternate) in the Settings, you can see the alternate pattern clearly.
The motion aftereffect (MAE) is a visual illusion that after watching a moving stimulus for some time, stationary objects appear to move in the opposite direction. For example, if you stair the stimulus for 30 seconds, and then click stop all button, you will see a motion toward the opposite direction.
Current version is one variation of the MAE. You should see an combined effect of the stimuli, when you stop the stimuli.
This project is licensed under the Creative Commons Attribution-NonCommercial 4.0 International License by Guang Yang (Spike).
Anstis, S., Verstraten, F. A., & Mather, G. (1998). The motion aftereffect. Trends in cognitive sciences, 2(3), 111-117.